- 2006-2011, Founding Director, MRC Centre for Regenerative Medicine, University of Edinburgh;
- 2005-2006, Professor of Reproductive Science, University of Edinburgh;
- 2000 -2005, Head of Department, Gene Expression and Development, Roslin Institute;
- 1973-2005, Principal Investigator, Roslin Institute;
- 1971-1973, Postdoc, ARC Unit of Reproductive Physiology and Biochemistry, Cambridge. The first methods were developed for the deep freeze storage of mammalian embryos.
- 1967-1971, PhD Darwin College, Cambridge Deep Freeze Preservation of Boar Semen.